Festuca of China

Xiang Chen, S. G. Aiken, and M. J. Dallwitz


Festuca leptopogon Stapf J.D.Hooker. Fl. Brit. Ind. 7: 354. 1897.

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Festuca subulata var. leptopogon (Stapf) Saint-Yves. 1928. Festuca takasagoensis Ohwi. 1933. Subg. Subulatae (Tzvelev) E. B. Alexeev, Parviglumae S. Aiken & X.Chen sect. nov.

Vegetative morphology. Plants softly erect. Plants loosely tufted. Plants clumped loosely. Plants with non-flowering shoots less common than flowering shoots, or non-flowering shoots rare or absent. Plants with leaves along the stems (usually). Plants with tillering leaves 1/4 to 1/3 the length of the culms. Plants with shoots extravaginal. Rhizomes present. Rhizomes short. Culms (30–)60–120 cm tall. Culms 1.1–1.3 mm wide. Culms geniculate, or strict, erect (usually). Culms internodes glabrous. Culms nodes 3–5. Uppermost culms nodes in lower 1/3 of the culm, or in mid 1/3 of the culm. Sheaths shorter than the internode. Sheaths glabrous. Sheaths open more than half their length. Sheaths not conspicuous at the base of the plant. Sheaths splitting between the veins. Auricles absent. Ligules (0.5–)1–2 mm long. Ligules glabrous. Ligules membranous. Ligules margins without cilia. Ligules apices erose. Leaf blades more or less lax. Leaf blades linear. Leaf blades flat (tender). Leaf blades adaxial lateral longitudinal ridges indistinctly raised. Leaf blades 8–15(–30) cm long. Leaf blades 3–8 mm wide. Leaf blades adaxial surfaces with trichomes. Leaf blades adaxial surfaces scabrous. Leaf blades abaxial surfaces glabrous, or with trichomes. Leaf blades abaxial surfaces scabrous (scaberulous). Leaf blades margins smooth, or scabrous. Flag leaves blades (5–)9–12 cm long. Flag leaves blades nearly equal to the sheath (usually). Veins 18–22. Adaxial to abaxial sclerenchyma strands present. Abaxial sclerenchyma poorly developed.

Floral morphology. Inflorescences narrowly lanceolate, or lanceolate. Inflorescences loosely open. Inflorescences (10–)15–30 cm long. Inflorescences branches 6–15 cm long. Inflorescences lowest internodes 4–10 cm long. Inflorescences branches at the lowest nodes 1 (usually), or 2. Inflorescences branches flexible. 12–13 fertile spikelets on the longest branches. Inflorescences branches angular in cross section, or flattened and ribbon-like. Inflorescences branches scabrous on the angles. Inflorescences branches vestiture sparse, or moderate. Pedicels 1–7 mm long. Spikelets lower third of the branches without spikelets, or lower half of the branches without spikelets. Spikelets 7–8 mm long. Spikelets 1.5–2.5 mm wide. Spikelets greenish. Spikelets narrowly lanceolate, or lanceolate. Florets 2–3. Glumes herbaceous. Glumes glabrous, or with trichomes. Glumes surfaces scabrous. Glumes vestiture on the veins (midveins). Glumes margins not ciliate. Glumes margins broadly membranous. Glumes apices accuminate, or acute. First glume lanceolate. First glume 1.5–3.2 mm long. First glume 1 nerved. First glume 0.7 as long as second glume. Second glume broadly lanceolate. Second glume 2.5–4(–5.5) mm long. Second glume 3 nerved. Second glume shorter than first lemma. Second glume 0.5–0.6 as long as the spikelets. Rachilla internodes 1–1.5 mm long. Rachilla internodes antrorsely scabrous. Rachilla internodes cylindrical. Rachilla internodes narrow. Rachilla internodes straight. Lemmas 6.5–7.5(–8) mm long. Lemmas lanceolate. Lemmas somewhat dorsally compressed. Lemmas veins 5. Lemmas with distinct veins in dorsal view. Lemmas glabrous. Lemmas upper margins narrowly membranous, or not membranous. Lemmas apices notched (distinctly). Lemmas apices accuminate. Lemmas apices awns 6–10(–15) mm long. Lemmas apices awns about as long as the lemmas, or much longer than the lemmas. Paleas 5 mm long. Paleas 0.5–0.7 mm apart. Paleas equal or subequal to the lemmas. Paleas keels glabrous (usually). Paleas inter-keel regions glabrous. Anthers 1–1.2(–1.6) mm long. Ovary apices hairy. Ovary hairs moderate, or dense. Fruits 3.5–4.2 mm long (immature).

Ecology. Flowering time May-Jul. Under forests of mountain slopes , in grasslandss or along streamsides; 2300–3900 m. Distribution. Northwestern China: Qinghai. Southwestern China: Sichuan, Yunnan, Guizhou, Xizang. Southern China: Taiwan.

Notes. In all our analyses this taxon aligned with other members of sect. Parviglumae. Lu (1992) had placed it in sect. Longiglumes.

Illustrations. • Herbarium specimen: PE: 1300041. Herbarium label reads: collected in Sichuan on slope, among shrubs. Note long flat leaves, loose panicle and narrow spikelets. • Type specimen of F. takasagoensis. Holotype of F. takasagoensis Ohwi at Herb. Univ. Kyoto, Ohwi. 2735. treated as a synonom of Festuca leptopogon Stapf. • Plant inflorescence, spikelet, and floret. Drawing from Fl. Ill. Pl. Prim. Sin. Gram. 1959. Inflorescence lanceolate, loosely open, (10-)15–30 cm long. Spikelets 7–8 mm long, 1.5–2.5 mm wide, greenish, lanceolate or narrowly lanceolate. Florets 2–3(usually). First glumes lanceolate, 1.5–3.2 mm long, 1 nerved, 0.7 as long as second glumes. Second glumes broadly lanceolate, 2.5–4(-5.5) mm long, 0.5–0.65 as long as the spikelets, 3 nerved, shorter than the first lemma. Lemmas apices apically cleft (distinctly). • Close up to the spikelet. Spikelet approximately 8 mm long, with relately short and narrow, 3 florets with awns longer than the body of the lemma. Spikelet from PE: 1300041. • Distribution map.


Cite this publication as: ‘Xiang Chen, S.G. Aiken, and M.J. Dallwitz (2002 onwards). Festuca of China: Descriptions, Illustrations, Identification, and Information Retrieval. Version: 4th February 2003. http://www.cdelta.ibcas.ac.cn’.

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